capital letter

noun
a letter of the alphabet that usually differs from its corresponding lowercase letter in form and height, as A, B, Q, and R as distinguished from a, b, q, and r : used as the initial letter of a proper name, the first word of a sentence, etc.
Also called capital.
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Based on the Random House Dictionary, © Random House, Inc. 2013.
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Etymonline
Word Origin & History

capital letter
late 14c.; see capital. So called because it is at the "head" of a sentence or word.
Online Etymology Dictionary, © 2010 Douglas Harper
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00:10
Capital letter is always a great word to know.
So is zedonk. Does it mean:
the offspring of a zebra and a donkey.
a children's mummer's parade, as on the Fourth of July, with prizes for the best costumes.
Example sentences
Usually the symbol is the combination of an initial capital letter followed by
  one or more small letters.
Type is measured by measuring a capital letter from the baseline to the top of
  the cap.
To indicate a capital letter, dot six is placed before the cell.
And having to resubmit the entire proposal, again and again, for one word that
  lacked a capital letter is ludicrous.
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