multiple factors

noun Genetics.
a series of two or more pairs of genes responsible for the development of complex, quantitative characters, as size or yield.

Origin:
1910–15

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World English Dictionary
multiple factors
 
pl n
genetics two or more genes that act as a unit, producing cumulative effects in the phenotype

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Example sentences
But if multiple factors are present and persist it can result in sustained aggression or even colony collapse.
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